CaliforniaHerps.com

A Guide to the Amphibians
and Reptiles of California


Panamint Rattlesnake - Crotalus stephensi

Klauber, 1930
Click on a picture for a larger view
Speckled Rattlesnakes Range MapOrange: Range of this species in California
Crotalus stephensi - Panamint Rattlesnake

Range of similar species in California:

RedCrotalus pyrrhus -
Southwestern Speckled Rattlesnake



Click on the map for a topographical view

Map with California County Names






observation link






Venomous and Potentially Dangerous!

Panamint Rattlesnake
Adult, Inyo County
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Adult, 6,300 feet elevation, White/Inyo Mountains, Inyo County Adult, Argus Range, Inyo County
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Adult, White/Inyo Mountains, Inyo County Adult, Inyo County Adult, Inyo County © Brad Alexander
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Adult, Kingston Mountains, San Bernardino County © Keith Condon Adult, southern Sierra Nevada,
Inyo County © David Miles
Adult, Inyo County
© John Stoklosa
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Adult, Death Valley, Inyo County © Joel Lingenfelter Adult, Inyo County © Richard Porter Adult, Inyo County © Ryan Sikola
rattlesnake Panamint Rattlesnake Panamint Rattlesnake
rattlesnake
Adult male, Mono County
© Adam G. Clause
Adult, San Bernardino County 
© Jacob Anderson
Adult, San Bernardino County 
© Jacob Anderson
Adult, Saline Valley, Inyo County
© Terry Goyan
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake rattlesnake
Adult, San Bernardino County © Kenny Elliott Adult, San Bernardino County
© Zachary Lim
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Comparisons of two adult Panamint Rattlesnakes found at the same spot in Inyo County. © Ryan Sikola Adult, western Nevada
© Brad Alexander
rattlesnake rattlesnake rattlesnake rattlesnake
Adult, Inyo County © Ryan Sikola Adult, Kern County
© Zeev Nitzan Ginsburg
Adult, San Bernardino County
© Zeev Nitzan Ginsburg
rattlesnake rattlesnake
Dark juvenile from San Bernardino County © Rob McLennan
rattlesnake Panamint Rattlesnake rattlesnake  
Adult, Inyo County © Rob McLennan Tail and Rattle

Comparison of the tails of
C. stephensi
and C. mitchellii
Adult, southern Sierra Nevada, southern Inyo County © Carl Brune

Some experienced herpers have looked at this photo and identified this snake as a Northern Pacific Rattlesnake, C. o. oreganus. Find more information about the controversy of identifying rattlesnakes from the southern Sierra Nevada Mountains region here.
 
       
Juveniles
Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake Panamint Rattlesnake
Juvenile, White/Inyo Mountains, Inyo County Juvenile, eastern Sierras, Inyo County
© Chris Morrison
Sub-adult, eastern Sierras, Inyo County © Chris Morrison
rattlesnake rattlesnake    
Subadult female, Mono County
© Adam G. Clause
Juvenile, Inyo County © Ryan Sikola    
       
California Kingsnake Predation
california kingsnake california kingsnake california kingsnake california kingsnake
california kingsnake california kingsnake    
Stacy Holt with Death Valley National Park sent me the above six photos which were taken on 8/28/13 by National Park Service Employees Drew Kaiser and Shannon Mazzei. Drew and Shannon saw the snakes struggling at around 11 AM in near Towne Pass. A California Kingsnake was wrapped tightly around a Panamint Rattlesnake and the snakes were barely moving. Disturbed by the onlookers, the kingsnake retreated under a nearby bush. The rattlesnake was dead by that time, and appears to be biting itself, but was described as biting onto the kingsnake before it died. The kingsnake probably returned to swallow the rattlesnake after the people left.
You can see other interesting wildlife sightings on the Death Valley National Park Facebook Page.
       
How to Tell the Difference Between Rattlesnakes and Gophersnakes
Gopher Snake Rattlesnake Comparison Sign sign Gopher Snake Rattlesnake Comparison Sign Gopher Snake Rattlesnake Comparison Sign
Harmless and beneficial gophersnakes are sometimes mistaken for dangerous rattlesnakes. Gophersnakes are often killed unnecessarily because of this confusion.
(It's also not necessary to kill every rattlesnake.)

It is easy to avoid this mistake by learning to tell the difference between the two families of snakes. The informational signs shown above can help to educate you about these differences. (Click to enlarge).

If you can't see enough detail on a snake to be sure it is not a rattlesnake or if you have any doubt that it is harmless, leave it alone.
You should never handle a snake unless you are absolutely sure that it is not dangerous.

   
Habitat
Panamint Rattlesnake Habitat Panamint Rattlesnake Habitat Panamint Rattlesnake Habitat Panamint Rattlesnake Habitat
Habitat, 6,300 feet elevation,
Inyo County
Habitat, 5,900 feet elevation, White/Inyo Mountains, Inyo County Habitat, Death Valley, Inyo County © Joel Lingenfelter

Panamint Rattlesnake Habitat      
Habitat, Inyo County
     
       
Sound and Video
Panamint Rattlesnake speaker icon


   
A Panamint Rattlesnake found on a road at night in Inyo County, rattles and crawls away. Click on the play button or the speaker to hear a rattlesnake rattling.
   
       
 
sign
rattlesnake sign sign
  California State Park warning sign.
Click the picture to see more rattlesnake signs.

 

Rattlesnakes are important members of the natural community. They will not attack, but if disturbed or cornered, they will defend themselves. Reasonable watchfulness should be sufficient to avoid snakebite. Give them distance and respect.

"Rattlesnakes are also among the most reasonable forms of dangerous wildlife: their first line of defense is to remain motionless; if you surprise them or cut off their retreat, they offer an audio warning; if you get too close, they head for cover. Venom is intended for prey so they're reluctant to bite, and 25 to 50 percent of all bites are dry - no venom is injected."  
Leslie Anthony. Snakebit: Confessions of a Herpetologist. Greystone Books, 2008.

Rattlesnakes are typically described as poisonous, but they are actually venomous.
A poisonous snake is one that is harmful to touch or eat. A venomous snake injects dangerous venom into its victim.

A bite from a rattlesnake can be extremely dangerous, but rattlesnakes should not be characterized as aggressive and vicious, striking and biting without provocation, as they are often shown. If rattlesnakes are given some space and enough time to escape to a safe place, they will usually just crawl away as fast as possible to avoid confrontation. Rattlesnakes will not strike without a reason: they will strike at a potential meal and they will defend themselves from anything they perceive as dangerous. They avoid striking and biting because it uses up their valuable supply of venom which they need to kill and digest their food.

Rattlesnakes are often portrayed with the body partly coiled, the tail rattling loudly, and the head raised up and ready to strike, but they do not need to coil up this way to strike and bite. This display is a warning not to come any closer. It's a defensive behavior that some rattlesnakes use when they sense that crawling away would put them in danger of attack.

Rattlesnakes do not always rattle a warning. Sometimes they rattle loudly to warn potential enemies of their presence, but other times they remain silent when they sense a threat, choosing to remain still to rely on their cryptic color and pattern to let them blend into their surroundings to hide from the threat. Making a noise in this situation risks advertising their presence. They also use their natural camouflage to hunt by sitting still, without rattling, trying to remain invisible as they wait for a warm-blooded prey animal to pass close enough to strike.

Description

Dangerously Venomous


A bite from this snake can cause death or serious illness or injury in humans that may require immediate medical care.

(Commonly called a "poisonous" snake to indicate that its bite is dangerous, but that is not correct. It should be called a "venomous" snake. A poisonous snake can harm you if you eat it. A venomous snake can harm you if it bites you.)

Size
Adults are 23-52 inches in length (58-132 cm) averaging 2 - 3 feet (61-91 cm).
Hatchlings are about 10 inches (25 cm).

Appearance
A long, heavy-bodied snake with a thin neck, a large triangular head, and a rattle on the end of the tail consisting of loose interlocking hollow segments.  A new rattle segment is added each time the skin is shed, which can be more than one time per year.
Pupils are elliptical.
Scales are keeled.

Bites on humans are potentially dangerous without immediate medical treatment.
Even a dead snake can bite and inject venom if the jaws reflexively open when they are touched.

Rattlesnakes are "pit vipers" which means they have two pits that are used to sense heat when hunting warm-blooded prey - with one pit on each side of the front of the head above the mouth.
Color and Pattern
Shows a great variety of body coloration which usually allows the snake to blend into its environment - tan, yellowish, orangish, gray, off-white, brown.
The body is marked with a pattern consisting of dark speckled banded markings, which can be vague or distinct.
A dark band or bands on the tail, but not usually alternating with light bands.
The ground color of the tail is generally the same as the body color, not contrasting sharply with it.
The last dark tail bands often seem to fuse together into one large black band just before the rattle.
Compare
with the tail and rattle of the similar species C. m. pyrrhus.
Young
Juveniles are born with only a silent button at the end of the tail.
Similar Snakes
Found in sympatry with other species of rattlesnakes, including Crotalus scutulatus and Crotalus cerastes. Can be differentiated from these species by color, pattern, and tail rings, and the lack of horns over the eyes.
Very similar to Crotalus mitchellii pyrrhus, which was once thought to be the same species. Information on differentiating the two species can be found here.

Life History and Behavior

Activity
Primarily nocturnal and crepuscular during periods of excessive daytime heat, but also active during daylight when the temperature is more moderate. Not active during cooler periods in Winter.
Fangs and Venom
Two large hollow movable fangs are located at the front of the upper jaw are folded backwards when not used. The fangs are connected to venom glands so that when the snake bites, the fangs swing forward rapidly to stab the prey and inject a toxic venom that quickly immobilizes the prey. A rattlesnake can control the amount of venom injected. The fangs can be replaced if broken.

Bites that inject venom into humans are potentially dangerous without immediate medical treatment.
Sometimes a rattlesnake bites but does not inject venom. These are called "dry bites." A dry bite may still require medical attention.
Even a dead snake can bite and inject venom if the jaws open and close reflexively when they are touched.

A bite from any kind of rattlesnake of any age or any size should be treated as a serious medical emergency, but the bite of a juvenile rattlesnake is not more dangerous than the bite of an adult.
Experts disagree whether or not juvenile rattlesnake venom is more potent than adult rattlesnake venom, but this does not really make much difference in the severity of a bite.
While adult rattlesnakes can control the amount of venom they inject depending on their needs (small animals need less venom, a defensive or warning bite may need no venom, etc.), it is often assumed that juvenile rattlesnakes do not have the same ability and that they always inject the full amount of venom they have available. Some studies show this is not true. There is also no proof that adult rattlesnakes are more likely than juveniles to bite without injecting venom when they are biting as a warning. Regardless of these things, adults have far more venom to inject than juveniles so the potential danger from the bite of an adult is significantly higher than the danger from the bite of a juvenile. Even when an adult does not inject the full amount of venom it has available, it most likely injects more venom than a juvenile would inject.

Venomous snakes are immune to the venom of their own species, so if a snake is bitten during interactions with other snakes of its species during territorial fights or during mating or if it accidentally bites itself, it will not suffer from the venom. However, they are not typically immune to the venom of other species of snakes.
Sound - The Rattle
When alarmed, a rattlesnake shakes its tail back and forth. The movement rubs the rattle segments together producing a buzzing sound which serves as a warning.
Diet and Feeding
Eats small mammals, lizards, and birds.

Heat sensing pits on the sides of the head help the snake to locate prey by their warmth.
Long, hollow, movable fangs connected to venom glands inject a very toxic venom which quickly immobilize prey.
The snake can control the amount of venom injected and the fangs are replaced if broken.

An ambush hunter, it may wait near lizard or rodent trails, striking at and releasing passing prey. The snake then follows the trail of the envenomated animal and swallows it whole. Prey is also found while the snake is actively moving.
Perception
rattlesnake perception

Click on this picture to see an illustrated interpretation of the various ways pit vipers (including rattlesnakes) perceive their prey, using their eyes, their sense of smell, their ability to detect vibrations, and their ability to sense heat.
© Frank Buchter
Reproduction
Rattlesnakes are ovoviviparous. The mother keeps her fertilized eggs inside her body and gives birth to living young.
Females probably start bearing young at three years of age and breed annually. (Klauber, 1982)
Mating occurs in the spring.
Two to 12 young are born in July or August. (Stebbins & McGinnis, 2013)


Male "Combat Dance"

Adult males engage in a ritual "Combat Dance" during the spring breeding season and at other times. Throughout much of history this activity was presumed to be a mating male and female instead of two competing males. Despite the common name, it is not combat as neither male is injured. And it's not a dance, it's essentially a wrestling match in which necks and forebodies are intertwined, with the stronger snake slamming the smaller one to the ground until the weaker snake leaves the area. Most bouts end in a draw.
"Certainly the presence of a female is not necessary to stimulate males to dance."
"Dancing is not restricted to a single season of the year."
(Klauber, 1982)
Habitat
Associated mostly with habitats composed of rocky outcrops and boulders, but also found in creosote bush and cactus deserts and open coniferous woodlands.

Geographical Range
Found in central eastern California, from approximately the Mojave River north along the east side of the Sierras into Nevada.

Traditional range maps for this snake, including my former maps, show it ranging west in the Mohave Desert in Kern County to the Tehachapi mountains, but I have found no museum records from this area farther south than about 20 miles south of Randsburg and no observations from the area on iNaturalist, so I no longer show it occurring in this area.

"Klauber (1930, 1936) suggested that C. m. stephensi formed a zone of intergradation with C. m. pyrrhus in the Mohave Desert (i.e., the Barstow–Ivanpah–Hoover Dam line). His primary evidence was the presence of an incomplete separation of the prenasal and rostral scales in some individuals he examined from this region. This condition, however, occurs infrequently throughout the distribution of C. mitchellii (Klauber, 1936, 1949, 1963), and hence its utility as a robust morphological indicator of intergradation between C. m. stephensi and C. m. pyhrrus is unresolved."

Full Species Range Map
Elevational Range
Sea level to 8,000 ft. (2,440 m).

Notes on Taxonomy
In a 2007 paper, * using molecular data, Douglas et al showed that this snake is a distinct species, not a subspecies of Crotalus mitchellii.


Alternate and Previous Names (Synonyms)

Crotalus stephensi - Panamint Rattlesnake (Douglas, Douglas, Schuett, Porras, Thomason 2007, Stebbins & McGinnis 2012)
Crotalus mitchellii stephensi - Panamint Rattlesnake (Stebbins 1966, 1985, 2003, Klauber 1982)
Crotalus mitchellii stephensi - Panamint Rattlesnake (Wright & Wright 1957)
Crotalus mitchellii stephensi (Stebbins 1954)
Crotalus mitchellii stephensi (Klauber, 1930)
Crotalus tigris - Tiger Rattlesnake) (Grinnell and Camp 1917)
Crotalus mitchellii (Cope, 1861)

Conservation Issues  (Conservation Status)
None
Taxonomy
Family Viperidae Vipers Crotalidae - Pitvipers
Genus Crotalus Rattlesnakes Linnaeus, 1758
Species


stephensi Panamint Rattlesnake Klauber, 1930
Original Description
Crotalus mitchellii - (Cope, 1861) - Proc. Acad. Nat. Sci. Philadelphia, Vol. 13, p. 293
Crotalus mitchellii stephensi - Klauber, 1930 - Trans. San Diego Soc. Nat. Hist., Vol. 6, No. 3, p. 108

from Original Description Citations for the Reptiles and Amphibians of North America © Ellin Beltz

Meaning of the Scientific Name
Crotalus - Greek - krotalon - a rattle - refers to the rattle on the tail
mitchellii
- honors Mitchell, S. Weir
stephensi - honors Stephens, Frank

from Scientific and Common Names of the Reptiles and Amphibians of North America - Explained © Ellin Beltz

Related or Similar California Snakes
C. pyrrhus - Southewstern Speckled Rattlesnake
C. s. scutulatus - Northern Mohave Rattlesnake
C. c. cerastes - Mojave Desert Sidewinder
C. o. lutosus - Great Basin Rattlesnake
C. o. oreganus - Northern Pacific Rattlesnake

More Information and References
California Department of Fish and Wildlife

Living With Rattlesnakes

California Department of Fish and Wildlife: Rattlesnakes in California

University of California: Rattlesnakes Management Guide

Florida Museum of Natural History: How to Safely Coexist With Snakes

The Tucson Herpetological Society: Living With Venomous Reptiles

Washington Department of Fish and Wildlife: Living With Snakes

Centers for Disease Control and Prevention: Venomous Snakes

Anapsid.org: Melissa Kaplan's Rattlesnake Information Page

Southwestern Field Herping Association: Venomous Snake Safety


Snake Bites


California Poison Control System (search for "rattlesnake bite")

UCI Health - What to do if you're bitten by a rattlesnake

Snakebite Safety! How to Effectively Avoid, Identify, and Treat a Snake Bite (Includes all of the U.S.A.)

Don't Use Snakebite Suction Devices :
Snakebite Suction Devices Don't Remove Venom: They Just Suck
(Sean P. Bush, MD - Annals Of Emergency Medicine 43:2 pages 187-188 February 2004.)

Bay Nature Magazine - Are Baby Rattlesnakes the Most Dangerous Biters?

The Amazing Story of Andy Cat - a very lucky pet cat who was bitten by a rattlesnake and survived, thanks to the smart actions of its owners.

Wickipedia List of Fatal Snake Bites in the United States

Stebbins, Robert C., and McGinnis, Samuel M.  Field Guide to Amphibians and Reptiles of California: Revised Edition (California Natural History Guides) University of California Press, 2012.

Stebbins, Robert C. California Amphibians and Reptiles. The University of California Press, 1972.

Flaxington, William C. Amphibians and Reptiles of California: Field Observations, Distribution, and Natural History. Fieldnotes Press, Anaheim, California, 2021.

Samuel M. McGinnis and Robert C. Stebbins. Peterson Field Guide to Western Reptiles & Amphibians. 4th Edition. Houghton Mifflin Harcourt Publishing Company, 2018.

Stebbins, Robert C. A Field Guide to Western Reptiles and Amphibians. 3rd Edition. Houghton Mifflin Company, 2003.

Behler, John L., and F. Wayne King. The Audubon Society Field Guide to North American Reptiles and Amphibians. Alfred A. Knopf, 1992.

Powell, Robert., Joseph T. Collins, and Errol D. Hooper Jr. A Key to Amphibians and Reptiles of the Continental United States and Canada. The University Press of Kansas, 1998.

Bartlett, R. D. & Patricia P. Bartlett. Guide and Reference to the Snakes of Western North America (North of Mexico) and Hawaii. University Press of Florida, 2009.

Bartlett, R. D. & Alan Tennant. Snakes of North America - Western Region. Gulf Publishing Co., 2000.

Brown, Philip R. A Field Guide to Snakes of California. Gulf Publishing Co., 1997.

Ernst, Carl H., Evelyn M. Ernst, & Robert M. Corker. Snakes of the United States and Canada. Smithsonian Institution Press, 2003.

Taylor, Emily. California Snakes and How to Find Them. Heyday, Berkeley, California. 2024.

Wright, Albert Hazen & Anna Allen Wright. Handbook of Snakes of the United States and Canada. Cornell University Press, 1957.

Ernst, Carl. H. Venomous Reptiles of North America. Smithsonian Institution Press, 1999.

Hayes, William K., Kent R. Beaman, Michael D. Cardwell, and Sean P. Bush, editors. The Biology of Rattlesnakes. Loma Linda University Press, 2009.

Hubbs, Brian R., & Brendan O'Connor. A Guide to the Rattlesnakes and other Venomous Serpents of the United States. Tricolor Books, 2011.

Klauber, Laurence M. Rattlesnakes. University of California Press. (Abridged from the 1956 two volume Rattlesnakes: Their Habits, Life Histories, and Influence on Mankind.) University of California Press, 1982.

Rubio, Manny. Rattlesnake - Portrait of a Predator. Smithsonian Institution Press, 1998.

Walls, Jerry G. Rattlesnakes: Their Natural History and Care. T. F. H. Publications, Inc., 1996.

* Douglas, Michael E., Marlis R. Douglas, Gordon W. Schuett, Louis W. Porras, and Blake L. Thomason. Genealogical Concordance between Mitochondrial and Nuclear DNAs Supports Species Recognition of the Panamint Rattlesnake (Crotalus mitchellii stephensi). Copeia, 2007 (4), pp. 920–932.

Joseph Grinnell and Charles Lewis Camp. A Distributional List of the Amphibians and Reptiles of California. University of California Publications in Zoology Vol. 17, No. 10, pp. 127-208. July 11, 1917.

Conservation Status

The following conservation status listings for this animal are taken from the April 2024 State of California Special Animals List and the April 2024 Federally Listed Endangered and Threatened Animals of California list (unless indicated otherwise below.) Both lists are produced by multiple agencies every year, and sometimes more than once per year, so the conservation status listing information found below might not be from the most recent lists. To make sure you are seeing the most recent listings, go to this California Department of Fish and Wildlife web page where you can search for and download both lists:
https://www.wildlife.ca.gov/Data/CNDDB/Plants-and-Animals.

A detailed explanation of the meaning of the status listing symbols can be found at the beginning of the two lists. For quick reference, I have included them on my Special Status Information page.

If no status is listed here, the animal is not included on either list. This most likely indicates that there are no serious conservation concerns for the animal. To find out more about an animal's status you can also go to the NatureServe and IUCN websites to check their rankings.

Check the current California Department of Fish and Wildlife sport fishing regulations to find out if this animal can be legally pursued and handled or collected with possession of a current fishing license. You can also look at the summary of the sport fishing regulations as they apply only to reptiles and amphibians that has been made for this website.

This animal is not included on the Special Animals List, which indicates that there are no significant conservation concerns for it in California.
Organization Status Listing  Notes
NatureServe Global Ranking
NatureServe State Ranking
U.S. Endangered Species Act (ESA) None
California Endangered Species Act (CESA) None
California Department of Fish and Wildlife None
Bureau of Land Management None
USDA Forest Service None
IUCN


 

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